Higher plants evolved multiple innate immune systems to combat pathogens present in their living environment.However, activation of the complex immune responses allocates nutrient resources into biosynthesis of various defense molecules, and this in turn often represses growth and development of the entire plant.Over the years, the significance of the effective coregulation of growth and immunity has been demonstrated with dwarf phenotypes of autoimmune mutant lines of model plants displaying constitutive defense responses (van Wersch et al.,2016).This tightly regulated balance, termed the growthimmunity trade-off, is an important objective in plant breeding, which ultimately aims at plant cultivars that are both highly resistant and efficient in terms of growth and yield (Ning et al., 2017).