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Acyl-protein thioesterase 1 (APT1) is considered to be a key enzyme of S-depalmitoylation.Recently,Kathayat et al.[1] revealed the existence of APT1 in mitochondria,which was previously supposed to locate at cytosol and Golgi apparatus [2].Moreover,APT1 mediates the occurrence of mitochondrial S-depalmitoylation.