牡丹花药壁发育为基本型。花药壁由6～7层细胞构成,外中层宿存且发生带状纤维加厚。绒毡层有腺质和变形型两种类型。花粉母细胞减数分裂为同时型,小孢子四分体为正四面体型。生殖细胞初期具有呈PAS正反应的细胞壁,脱离花粉壁后被一层称为“脂体冠”的结构包围。成熟花粉为2细胞型,此时的生殖细胞为裸细胞,并与营养核紧密相连,构成了雄性生殖单位(MGU)。 Peony anther wall development as the basic type. Anther wall consists of 6 to 7 layers of cells, the outer middle layer of ribbon fibers and thickened. There are two types of tapetum, glandular and deformable. Pollen mother cells meiosis into simultaneous, microspore tetrad is a regular tetrahedron. The germ cells initially have a PAS-positive cell wall that leaves the pollen wall and is surrounded by a structure called the “fat crown.” Mature pollen is a 2-cell type, in which case the germ cells are naked cells and are closely linked to the vegetative nucleus, forming the male reproductive unit (MGU).