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1991年召开的第十一届国际组织相容性会议提出了140种HLA定型探针,其中用于DQB1的25种,DPB1的26种。在1992年底1993年初,Marsh等在组织抗原、免疫基因以及人类免疫学三种杂志上公布了32种DPB1等位基因,19种DQB1多位基因。
The Eleventh International Symposium on Compatibility held in 1991 proposed 140 HLA-typed probes, of which 25 for DQB1 and 26 for DPB1. At the end of 1992 and the beginning of 1993, Marsh et al. Published 32 DPB1 alleles and 19 DQB1 multi-loci genes in three types of tissue antigens, immune genes and human immunology.