Gibberellins (GA) are an important family of plant growth regulators, which are essential for many aspects of plant growth and development.In the GA signaling pathway, the action of GA is opposed by a group of DELLA family repressors, such as RGA.Although the mechanisms of action of the DELLA proteins have been studied in great detail, the effectors that act downstream of DELLA proteins and bring about GA-responsive growth and development remain largely unknown.