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It has been shown in our previous study that the Autographa californica nucleopolyhedrovirus (AcMNPV) ac76 gene is essential for both budded virion (BV) and occlusion-derived virion (ODV) development.More importantly, deletion ofac76 affects intranuclear microvesicle formation.However, the exact role by which ac76 affects the virion morphogenesis remains unknown.In this report, we characterized the expression, distribution, and topology of Ac76 to further understand the functional role of Ac76 in virion morphogenesis.